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Gramma, a Galápagos tortoise who was one of the oldest and most beloved residents of the San Diego Zoo, has died at an estimated age of 141. The zoo announced that she was humanely euthanized on Friday following a significant decline in her health and quality of life.
According to zoo officials, Gramma had been suffering from mobility issues and other age-related ailments that caused considerable discomfort. Dr. Nadine Lamberski, chief conservation and wildlife health officer for the San Diego Zoo Wildlife Alliance, stated that the animal care team made the difficult decision after exhausting all other options. She described Gramma as an "iconic and cherished individual" who connected generations of visitors to wildlife.
Gramma's history at the zoo dates back nearly a century. She was part of a group of tortoises brought to San Diego in 1928, donated by the crew of the U.S. Navy submarine tender USS Kanawha. Because she was already an adult upon her arrival, her exact age was always an estimate, but she was considered the zoo's longest-standing resident. Her remarkable longevity made her a symbol of the institution and a favorite among staff and guests, known for her affinity for her favorite foods like romaine lettuce and hibiscus flowers.
The loss of Gramma highlights the conservation challenges facing her species. Galápagos tortoises are listed as an endangered species, with their wild populations severely diminished over the last two centuries due to habitat loss and hunting. These giant tortoises are among the longest-living vertebrates on Earth, with some individuals known to live well over 150 years. Gramma lived through significant historical events and served as an ambassador for her species, inspiring conservation efforts. Her passing marks the end of an era for the San Diego Zoo, which she called home for 97 years.
